A nuclear tug-of-war to occupy the limited apical space after double disruption of deformation and dispersion pathways. Live imaging of kuk MT- embryos expressing RFP:NLS (false colored green) and Spider: GFP (magenta) zoomed to show tug-of-war behaviors. Each frame is acquired at every 15 s and video is played at 20 frames per second. Scale bar = 5 μm.